Mark studied ecology and conservation at Sparsholt College before completing BSc and MSc research degrees in ecology and biology at the University of Portsmouth. His research interests are in ecosystem function, trophic ecology, biogeochemistry and also visual communication and cognition of fishes. Mark has worked with Great White sharks, cetaceans, weird wood-eating molluscs and predatory flatworms and has described new Antarctic species from the deep sea. He is also a former public aquarium curator, author and aquarist with over 25 years experience.
